In the ovule or megasporangium, during the process of megasporogenesis, a hypodermal cell functions as a archesporium cell and becomes more prominent because of large size, denser cytoplasm and large nucleus. In tenuinucellate ovules, archesporial cell directly functions as megaspore mother cell while in crassinucellate ovules, archesporial cell divides periclinally to form an outer parietal cell and inner primary sporogenous cell that functions as megaspore mother cell.